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Curriculum
Vitae of Harvey B. Lillywhite May 1989
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Snake-A-Way
is the world's only EPA approved, university tested, patented snake
repellent. It has been proven effective against both poisonous and
non-poisonous snakes. When used and applied as directed it is
safe to human, animal and plant life. It has been universally
approved and endorsed by naturalist, conservationist and ecologists.
Natural
History Magazine 4/95 Vol. 104, no. 4
THE SERPENTS TONGUE by Kurt Schwenk
Aside
from the other senses, the most important and exquisitely sensitive
mode of the snake is the Jacobson's organ. In a world textured with
chemical clues, the snakes primary guide is its tongue. Recent
studies have demonstrated that the snake is always utilizing its
forked tongue to determine much about its surroundings. "Tongue
flicking delivers chemical particles into the mouth that makes its
way up through palate openings and into the Jacobson's organ,
stimulating sensory cells." It is similar to smell but
different and distinct." Like paired eyes or ears, that a snake
does not posses, a snakes forked tongue delivers a form of stereo
smell. It gives the snake the ability to sense the presence of their
enemies or prey. Temperature, humidity, acidity levels and prey
detection are just a few of the examples of the ability inherent in
this unique sensory receptor.
University
of Florida - Gainesville, FL
Summary
Test Animals: Snakes (various species)
Source: Various (private collections; wild / caught)
Date Animals Received Various June 1 - August 24
Date Tests Started: June 13, 1989
Date Tests Completed: October 2, 1989
Temperature & Humidity of Animal Room: 21+1*C: 50-65*RH
Behavioral
tests involved: (1) a two way choice situation within a Y-shaped
enclosure (Y) test, and (2) the use of space within a circular
enclosure in which one quarter of the area was bounded by a strip of
repellent (circle test). Snake-A-Way is shown to repel snakes with
efficacy that varies from 100% to 17% in the different taxa. The
nine venomous species tested (particularly rattlesnakes) exhibited
consistently high avoidance of the repellent, with the exception of
cottonmouth. The other snakes showed high avoidance of repellent,
except for rat snakes. Some individuals of all species demonstrated
overt avoidance of repellent. Collectively 83% of all snakes tested
avoided the strips of repellent used in the Y tests. Data for three
species of snakes subjected to circle tests corroborate the findings
from the Y tests. Thus, Snake-A-Way appears to be potent repellent,
however, snakes of all species may cross the repellent in unusual
circumstances. These circumstances are probably rare in nature,
however, and use of Snake-A-Way can be expected to reduce occurrence
of snakes in "protected" areas. Effectiveness tests were
conducted by Dr. Harvey Lilywhite, Professor of Reptology,
University of Florida. Dr. Lilywhite is internationally recognized
as one of the world's leading research repotoligst. Over ten years
of independent research and follow up tests were also conducted by
Dr. T's Nature Products, Inc. The effectiveness of the repellent is
demonstrated by the following experiments performed with various
species and the product has been found to have a rate of efficacy
that varies from 17% to 100% depending on the individual species.
87.5%
Agkistrodon contortirix, copperhead
21% Agkistrodon piscivorus, water moccasin
75% Coluber constrictor, black racer
91% Crotalus adamanteus, eastern diamondback rattle
91% Crotalus atrox, western diamondback rattlesnake
91% Crotalus, horridus, timber rattlesnake
91% Crotalus, lepidus, rock rattlesnake
91% Crotalus, ruber, red diamond rattlesnake
91% Crotalus, viridis, pacific rattlesnake
17% Elphe guttata, corn or red rat snake
47% Elaphe obsoleta, yellow rat snake
100% Heterodon platyrhinos, eastern hognose snake
100% Micrurus fulvivus, eastern hognose snake
50% Python fulvivus, eastern coral snake
100% Thamnophis marcianus, checkered garter snake
In
as much as the normal behavior of snakes is to move slowly in a
deliberate and investigatory manner, the experiments were devised to
test the snakes responses to the repellent in as near to normal
conditions as possible. The granules should be spread in a strip
10-30 centimeters wide, although generally the wider the strip the
better. In this manner, an odor barrier is provided that snakes are
reluctant to cross. Additionally, a smaller amount of the product
may be scattered in a diffuse or non-continuous manner just beyond
the strip, on the side away from the area to be protected. This
serves to lessen snakes wandering beyond the strip. After 2 1/2
months, a group of 16 snakes was again re-tested. In this test the
branches of the Y were reversed so that the repellent strip was at
the branch opposite from that in the previous tests. Moreover, the
repellent used was five weeks old (i.e., it had been on the ground,
outside, but protected from heavy rainfall). Test were conducted as
described from the primary test above. Eleven snakes exhibited overt
avoidance of the moving across the repellent strip while rubbing the
head against the ground, seemingly attempting to escape from the
repellent. Two of the snakes which crossed the repellent exhibited
avoidance behavior first, eventually moving across the repellent
strip while rubbing the head against the ground, seemingly
attempting to escape from the repellent. Repelled snakes showed no
hesitation to enter the opposite (being) branch of the Y.
The principle conclusions from this study are as follows: 1)
Snake-A-Way repels snakes. 2) The efficacy of the repellent varies
with taxon. 3) Circumstances in which individual snakes of all
species tested will cross a strip of repellent, so the product
cannot be regarded as an absolute barrier to these reptiles, 4) Use
of the product can be expected to reduce significantly the
occurrence of snakes within areas that are "protected" by
recommended use of the product.
Copperheads were strongly repelled by Snake-A-Way, while the related
cottonmouths were not. This result is interesting insofar as
cottonmouths inhabit swamps and readily feed on carrion. It is
possible that the sulfurous smell of the repellent is normally
encountered by the latter species, either in parts of its natural
habitat or during carrion feeding, or both.
Dr. Harvey Lillywhite,
Professor of Reptology, University of Florida conducted effectiveness tests. Dr.
Lillywhite is internationally recognized as one of the worlds leading research
reptologists. Dr. T's Nature Products, Inc., also conducted over ten years of
independent research and follow-up tests.
Address:
Department of Zoology University of Florida Gainesville, FL 32611
Telephone: (904) 392-1101
Date Of Birth: 1 December 1943
(Arizona, USA)
Marital Status: Married; 2 Children
Education:
PHD (Zoology: Physiological Ecology) UCLA, 1970 George A. Bartholomew, Committee
Chairman.
MA (Zoology: Physiological Ecology) UCLA, 1967
BA (Zoology: with honors) University of California Riverside, 1966
Professional
Experience:
1985 to present Professor, Department of Zoology,
University of Florida, Gainesville
January, February 1988 Research Expedition, central
Philippines (study and collecting of marine snakes).
March, April 1986 Research Expedition, central
Philippines (study and collecting of marine snakes). Funded by NIH.
1984 to 1985 Associate Professor, Department of Zoology,
University of Florida, Gainesville.
January, February 1985 Research Expedition, Visayas,
Philippines (Study and collecting of aquatic snakes). Funded by NIH.
1982 to 1984 Professor, Departments of Physiology and
Cell Biology and Systematic and Ecology, University of Kansas, Lawrence. August
1979 to
August 1980 Visiting Scientist, (sabbatical leave),
Physiological Research Laboratory, Scripps Institution of Oceanography,
University of California at San Diego. Funded by NIH.
June 1976 to June 1982 Associate Professor, Departments
of Physiology and Cell Biology and Systematic and Ecology, University of Kansas,
Lawrence.
June 1975 to June 1976 Visiting Lecturer and Visiting
Scientist, Department of Zoology, Monash University, Clayton, Victoria,
Australia.
July 1976 Research expedition, Papua New Guinea (study
and collecting of sea snakes. Funded by Australian Research Grants committee.
October 1971 to June 1976 Assistant Professor,
Physiology and cell Biology, University of Kansas, Lawrence.
Summer 1968 Training in development of miniature
telemetry systems: Biocom, Inc, Culver City, California.
October 1966 to June 1967 Blood Technician, veterans
Administration Hospital, West Los Angeles.
Teaching Experience:
University of Florida:
1. Animal Physiology
2. Reptilian Structure and Function
3. Integrated Principles of Biology
University of Kansas:
1. Biology of Organisms (undergraduate core
requirement)
2. Comparative Animal Physiology
3. Mammalian Physiology (team participant)
4. Respiration Physiology (team participant)
5. Human Physiology
6. Herpetology (team participant)
7. Ecology and physiology graduate seminars; research supervision
Monash University:
1. Physiology Ecology
2. Honors Student supervision
University of California, Los Angeles:
1. Animal Physiology (head teaching assistant)
2. Ecology and Evolution (teaching assistant)
3. Natural History of Animals(teaching assistant)
Professional Affiliations:
American Society of Zoologists; American Physiological
Society; American society of Ichthyologists and Herpetologists; American
Association for the advancement of Science; Ecological Society of America, Sigma
XI, and Phi Beta Kappa.
Other Affiliations:
Wilderness Society; Desert protective Council; National
Geographic Society; League of Conservation voters; The nature Conservancy; Union
of Concerned Scientists; National Parks and Conservation Association; Cultural
Survival; Word Wildlife Fund; Sierra Club legal Defense fund; Natural resources
Defense Council.
Publications
1968.
Lillywhite, H. & Maderson, P.F.A Histological changes in the epidermis of
the subdigital lamellae of Anolis carolinensis during the shedding cycle.
J.Morph. 125:379-402
1969. Lillywhite, H. Miniature
telemetry systems. Application Notes, Biocom Memo 1007. Biocom, Inc., Culver
City, California. (Technical Note)
1970. Lillywhite, H. Behavioral
temperature regulation in the bullfrog, Rana catesbeina. Copeia 1970:158-168.
1971. Lillywhite, H. Thermal
modulation of cutaneous mucus discharge as a determinant of evaporative water
loss in the frog, Rana catesbeina. Z. vergl. Physiology 73:84-104.
1971. Lillywhite, H. Temperature
selection by the bullfrog, Rana catesbeina. Comp. Biochem. Physiol,.
40A:213-227.
1973. Lillywhite, H., Light, P. &
Chelgren, P. The role of behavioral thermoregulation in the growth energetics of
the toad, Bufo boreas. Ecology 54:375-383.
1973. Lillywhite, H. (Review of) The
temperature and Water relations of Reptiles, by J. L. Clodseley-Thompson.
J.Syst.Zool. 22:201-202.
1974. Lillywhite, H. & Light, P.
Water movement over toad skin: functional role of epidermal sculpturing. Copeia
1974:257-258.
1974. Lillywhite, H. & North, F.
Perching behavior of Sceloporous occidentalls in recently burned chaparral.
Copeia 1974:984-986.
1974. Lillywhite, H. How frogs
regulate their body temperature. Environment Southwest No. 465, pp. 3-6.
1974. Seibert, E., Lillywhite, H.
& Wassersug, R. Cranial co- ossification in frogs: relationship to rate of
evaporative water-loss. Physiological Zoology 47:261-265.
1975. Lillywhite, H. & Light, P.
A comparative study of integumentary mucous secretions in amphibians.
1976. Lillywhite, H. Abundance and
diversity of lizards in a herbicide-treated environment. Am. Philos. Soc. Year
book 1975: 342-344.
1976. Seymour, R. & Lillywhite,
H. Blood pressure in snakes from different habitats. Nature 264:664-666.
1977. Lillywhite, H. Friedman, G.
& Ford, N. color matching and perch selection by lizards in recently burned
chaparral. Copeia 1977:115-121.
1977. Baldwin, J., Friedman, G.,
& Lillywhite, H. Adaptations to temporary muscle anoxia in anurans:
activities of glycolytic enzymes in muscles from species differing their ability
to produce lactate during exercise. Australia J. Zool. 25:15-18.
1977. Lillywhite, H. Effects of
chaparral conversion on small vertebrates in southern California. Biol. Conserv.
11:171-184
1978. Lillywhite, H. & Seymour,
R. Regulation of arterial blood pressure in Australian tiger snakes. J.exp.
Biol. 75:65-79.
1979. Lillywhite, H. (Review of)
Behavior and Neurology of Lizards, edited by N. Greenberg and P.D. MacLean.
Copeia 1978:558-560.
1979. Johnson, R.N. & Lillywhite,
H. digestive efficiency of the omnivorous lizard Klauberina riversiana. Copeia
1979:431-437.
1980. Lillywhite, H. Behavioral
thermoregulation in Australian elapid snakes. Copeia 1980:452-458.
1980. Roberts, J. & Lillywhite,
H. Lipid barrier to water exchange in epidermis of reptiles. Science
207:1077-1079.
1980. North, F. & Lillywhite, H.
The function of burrow turrets in a gregariously nesting bee. Southwestern
Naturalist 25:373-378.
1981. Gibbons, J.R.H. &
Lillywhite, H. Ecological segregation, color matching and speciation in lizards
of the Amphibolurus decresli species complex (Lacertilia: Agamidae). Ecology
62:1573-1584.
1981. Lillywhite, H. & smith, L.
Haemodynamic responses to haemorrhage in the snake, Elaphe obsoleta. J. Exp.
Biol. 94:275-283.
1982. Lillywhite, H. Tracking as an
aid in ecological studies of snakes. In N.J. Scott (ed.), Herpetological
Communities: a Symposium of the Society for the study of Reptiles and Amphibians
and the Herpetologist's League, August 1977. U.S. fish & Wildlife Research
Report 13. Pp. 181-191.
1982. Lillywhite, H. Cannibalistic
carrion ingestion by the rattlesnake, Crotalus virdis. J. Herpetol. 16:95.
1982. Lillywhite, H. & Maderson,
P.F.A, Skin Structure and Permeability. In C. Gans and F.H. Pough (eds.),
Biology of the Reptilia, Vol. 12, (Physiological Ecology). Academic Press, New
York & London. Pp. 397-442.
1983. Lillywhite, H. & Pough, F.H.
Control of arterial pressure in aquatic sea snakes. Amer. J. Physiol.
244:R66-73.
1983. Lillywhite, H., Ackerman, R.
& Palacios, L. Cardiorespiratory responses of snakes to experimental
hemorrhage. J. Comp. Physiol. 152:59-65.
1983. Roberts, J. & Lillywhite,
H. Lipids and the permeability of epidermis from snakes. J. Exp. Zool. 228:1-9.
1984. Pough, F. & Lillywhite, H.
Blood volume and blood oxygen capacity of sea snakes. Physiol. Zool. 57:32-39.
1984. Lillywhite, H. & Ackerman,
R. Hydrostatic pressure, shell compliance and permeability to water vapor in
flexible-shelled eggs of the colubrid snake Elaphe obsoleta. In Seymour, R.
(ed.), Respiration and Metabolism of Embryonic Vertebrates. Pp. 121-135.
1984. Lillywhite, H. Trailing
movements and sexual behavior in Coluber constrictor. J. Herpetol. 19:306-308.
1985. Lillywhite, H. Behavioral
control of arterial pressure in snakes. Physiol. Zool. 58:159-165.
1985. Lillywhite, H. & Gallagher,
K. Hemodynamic adjustments to head-up posture in the partly arboreal snake,
Elaphe obsoleta. J. Exp.Zool. 235:325-334.
1985. Lillywhite, H. B. Postural
edema and blood pooling in snakes. Physiol. Zool. 58:759-766.
1986. Smits, A. W. & Lillywhite,
H. Effects of hyperkalemia on thermoregulartory and feeding behaviors of the
lizard Sauromalus hispidus. Copeia 1986:518-520.
1987. Lillywhite, H. Circulatory
adaptations of snakes to gravity. Amer. Zool. 27:81-95.
1987. Lillywhite, H. & burggren,
W. Introduction to the symposium on cardiovascular adaptations in reptiles.
Amer. Zool. 27:3-4.
1987. Lillywhite, H. Temperatures,
Energetics and Physiological Ecology. In R. Seigel, J. Collins, and S. Novack
(eds.), Snakes: Ecology and Evolutionary Biology. MacMillan Pub. Co., New York
Pp. 422-477.
1987. Lillywhite, H. B. & Stein,
B. R. Surface sculpturing and water retention of elephant skin. J. Zool. (Lond.)
211:727-734.
1987. Lillywhite, H. Snakes under
pressure. Natural History 96:59-67. (reprinted in Science Horizons YearBook 1988
and Science Annual 1989, Franklin Watts, Grolier Enterprises. Danbury CT)
1988. Lillywhite, H. B. &
Maderson, P. F. A. The structure and permeability of integument. Amer.Zool.
28:945-962.
1988. Donald, J. A., & Lillywhite,
H. B. Adrengic innervation of the large arteries and veins of the semi-arboreal
rattlesnake, Elaphe Obsoleta. J. Morphol. 198:25-31.
1988. Lillywhite H., Smits, A. W.
& Feder, M. E. Body Fluid volumes in the aquatic snake, Acrochordus
Granulatus. J. Herpetol. 22:434-438.
1988. Lillywhite, H. B. (Review of)
Biology of the Reptillia, Volume 16, Ecology B, Defense and Life History. Copeia
1988:1102-1104.
1988. Lillywhite, H. B. Snakes, blood
circulation and gravity. Scientific American 256:92-98.
1989. Kattan, Ga. & Lillywhite,
H. B. Humidity acclimation and skin permeability in the lizard, Anolls
carolinensis. (in press, Physiol. Zool.)
1989. Donald, J. A. & Lillywhite,
H. B. VIP-immunoreactive nerves in the pulmonary vasculature of the aquatic file
snake, Acrochordus granulatus. Cell & Tissue Research 255:585-588.
1989. Donald, J. A. & Lillywhite,
H. B. Adrenergic nerves and 5-HT-containing cells in the pulmonary vasculature
of the aquatic file snake, Acrochordus granulatus. Cell & Tissue Research
256:113-118.
1989. Lillywhite, H.B. Unusual
shedding behaviors in the aquatic snake, Acrochordus grantulatus. Copeia 1989
(in press).
1989. Lillywhite, H. B. & Donald,
J.A. Pulmonary blood flow regulation in an aquatic snake. Science (in press).
Responses of snakes to odors are known to be important
in several contexts of their behavior. Such as locating prey and responding to
conspecifics (reviewed in Ford, 1986). Snakes are very sensitive to chemosenory
input, which involves the vomeronasal system. The tongue of a snake picks up
odorants, which mediates its analysis (Burghardt, 1980; Gillingham & Clark,
1981: Ford & Low, 1984). The vomeronasal system is involved in the
detection, localization and identification of odors (Cowles & Phelen, 1958;
Hapern & Kkubie, 1983). Thus, responses of snakes to Dr. T's Snake-A-Way
presumably are mediated by the detection of odorant molecules via the tongue and
vomernasal system. Behavioral tests designed to establish the efficacy of the
product in repelling snake must take into account the means of chemosensory
detection. Visual, tactile and auditory stimuli presumably are not relevant to
the intended use of this product. Thus, behavioral test were designed such that
moving snakes exploring their surroundings with tongue flicks encountered the
Snake-A-Way product as they might in real field situations.
In use, the resulting granular product is spread on the
ground across an area where it is desired to prevent or discourage snakes from
entering. Preferably, the granules should be spread in a strip 10-30 centimeters
wide, although generally the wider the strip the better. In this manner, an odor
barrier is provided that snakes are reluctant to cross. Additionally, a smaller
amount of the product may be scattered in a diffuse or non-continuous manner
just beyond the strip, on the side away from the area to be protected. This
added scattering of the repellent serves to lessen the probability of occasional
snakes wandering beyond the strip.
Snakes Tested:
All of the tests were conducted in outdoor enclosures,
which approximated natural situation insofar, as was practical within
constraints of the experimental design. Most of the snakes used in the study
were either newly born or recently captured, so the data was not dependent on
the long-term captives. Attempt was made to maximize the numbers of both species
and individuals subjected to testing. The following snakes (scientific and
common names were tested for response to Snake-A-Way.
87.5% Agkistrodon contortirix, copperhead
21% Agkistrodon piscivorus, water moccasin
75% Coluber constrictor, black racer
91% Crotalus adamanteus, eastern diamondback rattlesnake
91% Crotalus atrox, western diamondback rattlesnake
91% Crotalus horridus, timber rattlesnake
91% Crotalus lepidus, rock rattlesnake
91% Crotalus ruber, red diamond rattlesnake
91% Crotalus viridis, pacific rattlesnake
17% Elaphe guttata, corn or red rat snake
47% Elaphe obsoleta, yellow rat snake
100% Heterodon platyrhinos, eastern hognose snake
100% Micrurus fulvius, eastern coral snake
50% Python molurus, Burmese python
100% Thamnophis marcianus, checkered garter snake
In testing, snakes reacted to the repellent and
exhibited overt signs of distress or avoidance while in close contact with the
pellets. In other case, snakes turning away from the repellent before coming
within direct contact (with tongue) were sometimes re-tested and fount to
consistently turn away from the repellent while at distances of 10-30 cm. These
latter observations are noted as "apparent avoidance".
The effectiveness of the repellent is demonstrated by
the following experiments performed with various species and the product has
been found to have a rate of efficacy that varies from 17% to 100% depending on
the individual species.
Snakes were re-tested at the end of one week, using
week-old repellent. All snakes selected the benign branch of the Y and exhibited
overt avoidance behavior. These same snakes were then re-tested within one hour
and again avoided the repellent (seven snakes exhibiting the overt behaviors).
The group of snakes was then re-tested once more, 21 hours following the last
test just described. Again, these snakes avoided the repellent with the
exception of one individual, which began rubbing its head on the ground, crawled
haphazardly, and eventually "tunneled" its way through the repellent
strip.
Table 1; Figure 1
|
Fraction
of Snakes Species Crossing Repellent
|
|
Agkistrodon
Contortrix (Copperhead)
|
1/8
|
|
Agkistrodon
Piscivorus (CottonMouth or Water Moccasin)
|
6/15
|
|
Crotalus
(All Species)
|
2/37
|
|
Elaphe
Guttata (Corn or Red Rat Snake)
|
10/24
|
|
Elaphe
Obsoleta (Yellow Rat Snake)
|
4/15
|
|
Python
Molurus (Burmese Python)
|
3/15
|
|
Thamnophis
Marcianus (Checkered Garter Snake)
|
0/25
|
|
All
Other Species
|
0/4
|
After 2 1/2 months, a
group of 16 snakes was again re-tested. In this test the branches of the Y were
"reversed" so that the repellent strip was at the branch opposite from
that in the previous tests. Moreover, the repellent used was five weeks old
(i.e., it had been on the ground, outside, but protected from heavy rainfall).
Tests were conducted as described for the primary tests above. Eleven snakes
exhibited overt avoidance of the repellent, and four snakes crossed the
repellent. Two of the snakes which crossed the repellent exhibited avoidance
behavior first, eventually moving across the repellent strip while rubbing the
head against the ground, seemingly attempting to escape from the repellent.
Repelled snakes showed no hesitation to enter the opposite (benign) branch of
the Y.
Efficacy and
Species Variation:
With respect to testing, Snake-A-Way was 100% effective
in repelling snakes of some species while much less effective in repelling
others (table 1; figure 1). It is important to note that some individuals of all
species tested observed overt avoidance of the repellent. Extrapolating from
these results, it is expected that a significant fraction of snakes encountering
a strip of repellent in real-use situations will be repelled. The number or
percentage of repelled snakes should vary as a function of the species involved.
Snake-A-Way had the most dramatic effects in repelling
garter snakes (Thamnophis Marcianus), pythons (Python Molurus), and most of the
venomous snakes, particularly rattlesnakes(Crotalus SPP). The behaviors of
garter snakes, pythons, and coral snakes suggest that the repellent chemicals
are most obnoxious to these species. Other snakes (e.g. Crotalus Atrox)were
repelled with equal consistency, however.
Garter snakes are widespread throughout virtually the
entire continental United States and are commonly encountered in many parts of
the country. Although a single species was tested, it seems reasonable to assume
that Snake-A-Way could be used with considerable effectiveness to repel garter
snakes belonging to the Genus Thamnophis. The product also repels pythons
strongly. The species tested is Asiatic and commonly kept as a pet by reptile
fanciers in this country. Data for pythons suggest that the product likely will
repel the two species of endemic boid snake that occur in the western United
States.
Snake-A-Way should be an effective repellent for
rattlesnakes, which are a potential concern in many parts of the country. The
avoidance behaviors exhibited by snakes representing six different species (and
most geographic regions of the U.S.) suggest that the repellent is effective for
the genus, hence rattlesnakes generally. These venomous reptiles are of
widespread geographic occurrence and are locally abundant in many regions.
Copperheads were strongly repelled by Snake-A-Way,
while the related cottonmouths were not. This result is interesting insofar as
cottonmouths inhabit swamps and readily feed on carrion. It is possible that the
sulfurous smell of the repellent is normally encountered by the latter species,
either in parts of its natural habitat or during carrion feeding, or both.
Coral snakes exhibited overt avoidance of the
repellent. Which suggests that the product may be generally effective against
elapid snakes.
Rat snakes (Genus Elaphe)were not strongly effected by
Snake-A-way, although some individuals overtly avoided the repellent. Thus, the
product is expected to provide only partial effectiveness in repelling these
species.
Conclusions and
Discussions
The principle conclusions from this study are as
follows.
(1) Snake-A-Way repels snakes
(2) the efficacy of repellent varies with taxon:
some species are repelled with remarkable
effectiveness, while others are not. However, all species tested exhibit some
degree of behavioral avoidance of the repellent.
(3) There are circumstances in which individual snake
of all species tested will cross a strip of repellent, so the product cannot be
regarded as an absolute barrier to these reptiles. Such situations are expected
to be rare in natural environments, however, and,
(4) Use of the product can be expected to reduce significantly the occurrence of
snakes within areas that are "protected" by recommended use of the
product.
In cases where snakes exhibited avoidance behaviors but
nonetheless crossed the repellent strip in various ways, it seems likely that a
diffuse scattering of Snake-A-Way pellets over the area beyond the strip would
have caused the snake to eventually leave the area. Such procedure might also
enhance the effectiveness of the product on substrates that are coarse-grained
or uneven. Considering all of the data, the fundamental conclusions regarding
efficacy remain the same. Snake-A-Way should significantly decrease snake
occurrences.
UNIVERSITY OF FLORIDA
Gainesville, FL 32611
Department of Zoology
233 Bartram Hall
904-392-1107
SUMMARY
Test Animal: Snakes (various species)
Source: Various (private collections; wild-caught
specimens from Alachua County, Florida; research breeding programs).
Date Animals Received: Various, June 1 through August
24.
Date Tests Started: June 13, 1989.
Date Tests Completed: October 2, 1989
Temperature & Humidity of Animal Room: 21+ 1oC: 50-65o RH.
Responses to Snake-A-Way, a putative snake repellent,
were examined in 160 snakes representing 15 species and four families.
Behavioral tests involved: (1) a two-way choice situation within a Y-shaped
enclosure (Y tests), and (2) the use of space within a circular enclosure in
which one quarter of the area was bounded by a strip of repellent (circle test).
Snake-A-Way is shown to repel snakes with efficacy that varies from 100% to 17%
in the different taxa. The nine venomous species tested (particularly
rattlesnakes) exhibited consistently high avoidance of the repellent, with the
exception of cottonmouths. The other snakes showed high avoidance of repellent,
except for rat snakes. Some individuals of all species demonstrated overt
avoidance of repellent. Collectively, 83% of all snakes tested avoided the
strips of repellent used in the Y tests. Data for three species of snakes
subjected to circle tests corroborate the findings from the Y tests. Thus,
Snake-A-Way appears to be a potent repellent, however, snakes of all species may
cross the repellent in unusual circumstances. These circumstances are probably
rare in nature, however, and use of Snake-A-Way can be expected to reduce the
occurrence of snakes in "protected" areas.
Excerpts from The
University of Florida (EPA TESTING & PROTOCOL)